Cox was only 16 months old when Storey won her first Paralympic medal
at the 1992 Games in Barcelona, where she competed as a swimmer.
Cox had a stroke aged 23, which led to her being diagnosed with multiple sclerosis.
Storey,
a 12-time gold medallist who won the C4-5 time trial at London 2012,
completed the 500m race in 37.068 seconds to take the lead.
However,
Cox triumphed in 34.598 seconds, with China's Zhou Jufang taking silver
in 36.004 and compatriot Ruan Jianping bronze in 36.557.
Cox said: "I have worked so hard, my legs were tired and I was mentally exhausted from yesterday."
Storey said: "I'm a pursuiter - it should be a sprinter winning it.
"Fourth is the worst place to be, but even if I'd gone as quick as I did in London I'd still have been fourth."
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